Research Institute of Forests and Rangelands
The Iranian Journal of Botany
1029-788X
2383-1685
15
1
2009
06
22
EXPERIMENTAL TAXONOMY OF THE GENUS SALIX L. (SALICACEAE) IN IRAN
3
20
EN
Ali Asghar
Maassoumi
0000-0002-0134-4986
Professor / Department of Botany, Research Institute of Forests and Rangelands P. O. Box 13185-116, Tehran-Iran.
maassoumi@rifr-ac.ir
The genus <em>Salix</em> in Iran is revised. In this study a living collection of the species in Iran was prepared and used in taxonomic and experimental studies. As a result of this study the total number of species in Iran was increased from 12 to 31. Also, 7 hybrids were recognized in Iran.. The species <em>S. daviesii, S. dinsmorei, S. persica </em>and<em> S. australior</em> were excluded from the synonymy of their related species, <em>S. acmophylla and S. excelsa</em>. Comparing the species in natural habitats and in cultivation several cryptic characters were reevaluated for preparing the key and descriptions. S. <em>zygostemon</em> is shown to be a biparental hibrid of <em>S. cinerea and S. elbursensis</em>. Several new records for Iran such as: <em>S. daviesii, S. persica, S. dinsmorei, S. australior, S. tetrasperma, S. fragilis, S. denticulata, S. pentandra, S. atrocinerea, S. cinerea, S. caprea, S. pedicellata </em>subsp<em>. pedicellata, S. pseudomedemii, S. caucasica, S. caspica, </em>and<em> S. fedtschenkoi </em>were recognized. <em>S. firouzkuhensis, S. lacus-tari, S. viridiformis </em>and<em> S. elymaitica </em>for the first time is described. For all species occurring in Iran key, short notes, habitat, distribution and general distribution are given. All living plants are in Karaj experimental station and the herbarium materials are preserved in the National Herbarium of Iran (TARI). <br />The genus <em>Salix</em> in Iran is revised. In this study a living collection of the species in Iran was prepared and used in taxonomic and experimental studies. As a result of this study the total number of species in Iran was increased from 12 to 31. Also, 7 hybrids were recognized in Iran.. The species <em>S. daviesii, S. dinsmorei, S. persica </em>and<em> S. australior</em> were excluded from the synonymy of their related species, <em>S. acmophylla and S. excelsa</em>. Comparing the species in natural habitats and in cultivation several cryptic characters were reevaluated for preparing the key and descriptions. S. <em>zygostemon</em> is shown to be a biparental hibrid of <em>S. cinerea and S. elbursensis</em>. Several new records for Iran such as: <em>S. daviesii, S. persica, S. dinsmorei, S. australior, S. tetrasperma, S. fragilis, S. denticulata, S. pentandra, S. atrocinerea, S. cinerea, S. caprea, S. pedicellata </em>subsp<em>. pedicellata, S. pseudomedemii, S. caucasica, S. caspica, </em>and<em> S. fedtschenkoi </em>were recognized. <em>S. firouzkuhensis, S. lacus-tari, S. viridiformis </em>and<em> S. elymaitica </em>for the first time is described. For all species occurring in Iran key, short notes, habitat, distribution and general distribution are given. All living plants are in Karaj experimental station and the herbarium materials are preserved in the National Herbarium of Iran (TARI).
Salix,Taxonomy,experimental studies,Iran,new species,distribution
https://ijb.areeo.ac.ir/article_102699.html
https://ijb.areeo.ac.ir/article_102699_7a5632807f73e6da660d888182ee9e32.pdf
Research Institute of Forests and Rangelands
The Iranian Journal of Botany
1029-788X
2383-1685
15
1
2009
06
22
NEW CHROMOSOME NUMBER AND KARYOTYPE ANALYSIS IN FOUR ASTRAGALUS L. (FABACEAE) SPECIES
21
26
EN
Masoud
Sheidai
Shahid Beheshti University G. C., Faculty of Biological Sciences, Tehran, Iran.
msheidai@yahoo.com
Farrokh
Gahremaninejad
0000-0001-5860-9976
ghahremaninejad@gmail.com
Karyotype studies were performed in four <em>Astragalus </em>species of <em>A. cornu-carpae</em>, <em>A. mucronifolius</em>, <em>A. talimansurensis</em> from the section <em>Leucocersis </em>and <em>A. crysostachys </em>from the section <em>Hymenostegis</em>. All four species showed 2n = 2x = 16 chromosome number and had metacentric and sub-metacentric chromosomes. The size of chromosomes varied from 0.75 mm in <em>A. talimansurensis</em> to 4.00 mm in <em>A. cornu-carpae</em>. The species studied differed significantly in the size of their chromosomes and in karyotype formulae indicating the role of quantitative change in the chromatin (DNA) material as well as chromosomes structural changes in the species diversification. The chromosome numbers of all four species studied are new to science. PCA ordination of the species showed their karyotype distinctness and that such data are of no use in the sectional delimitation in <em>Astragalus</em>.
Astragalus,Fabaceae,Karyotype,Iran
https://ijb.areeo.ac.ir/article_102701.html
https://ijb.areeo.ac.ir/article_102701_1925a0618ec140c1511b45643d8b4bc4.pdf
Research Institute of Forests and Rangelands
The Iranian Journal of Botany
1029-788X
2383-1685
15
1
2009
06
22
PHYLOGENY OF THE GENUS EREMURUS (ASPHODELACEAE) BASED ON MORPHOLOGICAL CHARACTERS IN THE FLORA IRANICA AREA
27
35
EN
Kosar
Naderi Safar
Department of Plant Biology / Faculty of Basic Sciences, Tarbiat Modares University.
Shahrokh
Kazempour Osaloo
Department of Plant Biology / Faculty of Basic Sciences, Tarbiat Modares University.
skosaloo@modares.ac.ir
Mehdi
Zarrei
Department of Biology , Faculty of Sciences, Shahed University of Tehran.
A phylogenetic analysis of <em>Eremurus</em> (<em>Asphodelaceae</em>)based on morphological data is presented. A total of twenty-five characters including 20 gross morphological, 4 palynological and 1 chromosomal characters were analyzed to reconstruct phylogenetic relationships for 24 taxa of <em>Eremurus</em> and <em>Trachyandra</em> <em>malosana</em> plus <em>Asphodelus tenuifolius</em> and two <em>Asphodeline</em> species as outgroups. Maximum parsimony approach as implemented in PAUP* using heuristic search and branch swapping option of tree bisection-reconnection were used for phylogenetic analyses. The analysis of characters with successive weighting using rescaled consistency index generated more resolved and supported trees than the analysis with equally weighting characters. The analyses showed that <em>T.</em> <em>malosana</em> is well allied with a clade of <em>Eremurus </em>species. Among <em>Eremurus</em> species analyzed, <em>E. persicus </em>and a subclade of<em> E. kopetdaghensis </em>and<em> E. luteus </em>comprise the basal most branches as sisters to the remainder. <em>Eremurus</em> subgenus <em>Henningia</em> with 15 taxa was paraphyletic; whereas, nine members of <em>Eremurus</em> subgenus <em>Eremurus</em> formed a well supported monophyletic group for which <em>E. furseorum</em> of the former subgenus was sister taxon. Sections<em> Henningia</em> (of subgenus <em>Henningia</em>) and <em>Eremurus</em> (of subgenus <em>Eremurus</em>) are also appeared to be non-monophyletic while the section <em>Ammolirion</em> (of subgenus <em>Eremurus</em>) is monophyletic. Medium size pollen grain, (25-50 µm) and chromosome number of 2n=14 are shared by <em>Eremurus</em> and <em>Trachyandra</em>. Three characters including non-branched inflorescence, basifixed stamens and ellipsoidal pollen grains are synapomorphies for the genus <em>Eremurus</em> solely. Tubular/campanulate flowers, incurved tepals and tepals with 3-5 nerves are synapomorphies for the subgenus <em>Eremurus</em>. Similarly exerted stamen is a synapomorphy for the subgenus <em>Eremurus</em> and <em>E. forseorum</em> (from subgenus <em>Henningia</em>). The tubular flowers (rather than campanulate) and the thinnest pollen exine are putative synapomorphies for the section <em>Ammolirion</em>. The remaining characters were evolved homoplasiously or ambiguously among these taxa.
Asphodelaceae,phylogeny,Morphology,Eremurus,Iran
https://ijb.areeo.ac.ir/article_102704.html
https://ijb.areeo.ac.ir/article_102704_d6cee0f26853093b0273362cc9ed9b6a.pdf
Research Institute of Forests and Rangelands
The Iranian Journal of Botany
1029-788X
2383-1685
15
1
2009
06
22
FOUR NEW SPECIES OF THE GENUS COUSINIA CASS. (ASTERACEAE) FROM IRAN
36
44
EN
Mostafa
Assadi
Research Institute of Forests and Rangelands.
masadi@rifr-ac.ir
Four new species of the genus <em>Cousinia</em> are described from Iran. <em>C. barezica</em> is based on a specimen collected from Barez mountain chains. The species belongs to the section <em>Spinuliferae</em> and related to <em>C. longifolia</em> and <em>C. sicigera</em>. It however, differs from both by having discolored leaves. <em>C. isfahanica</em> another new species from the section <em>Rigidissimae</em> is related <em>C. rigidissima</em> and <em>C. auranti</em> both from Afghanistan. This species has been based on a specimen collected from Isfahan, Semirom. <em>C. maassoumii</em> is described from adjacent localities of Delijan and Arak cities. It is from the section <em>Pugioniferae</em> and most closely related to <em>C. macroptera</em> but having different leaf and indumentum characters. <em>C. pseudocandolleana</em> is described based on specimen from Khorasan province. It is closely related to <em>C. candolleana</em> an endemic of Hamadan province, but differs from it by having stem and stem leaf characters.
Cousinia,Asteraceae,new species,Flora,Taxonomy,Iran
https://ijb.areeo.ac.ir/article_102714.html
https://ijb.areeo.ac.ir/article_102714_90ea69bf8138f931df5172cf4091025f.pdf
Research Institute of Forests and Rangelands
The Iranian Journal of Botany
1029-788X
2383-1685
15
1
2009
06
22
A NEW SPECIES OF ONOBRYCHIS SECT. AFGHANICAE (FABACEAE) FROM IRAN
45
50
EN
Hassan
Amirabadizadeh
Herbarium of Khorassan Research Center of Agricultural & Natural Resources
amirabadih@yahoo.com
Farangis
Ghanavati
Seed and Plant Improvement Institute, Karaj, Iran.
farangisghanavati@yahoo.com
Massoud
Abbassi
Herbarium of Khorassan Research Center of Agricultural & Natural Resources.
Massoud
Ranjbar
Department of Biology, Herbarium Division, University of Bu-Ali Sina.
ranjbar@baus.ac.ir
<em>Onobrychis iranensis</em> is described and illustrated as a new species from central and eastern Iran. The new species is related to <em>O. tavernieraefolia </em>with that it is compared here. Also the pollen morphology of these tow taxa were studied using scanning electron microscopy. The pollen grains are tri-colpate with reticulate ornamentation. Differences are observed in the size of pollen grains and the type of their aperture shape.
Hedysareae,new species,Onobrychis iranensis,Pollen morphology,Papilionaceae,Iran
https://ijb.areeo.ac.ir/article_102720.html
https://ijb.areeo.ac.ir/article_102720_2ad41beeae9588fb2261b2efda69fcf5.pdf
Research Institute of Forests and Rangelands
The Iranian Journal of Botany
1029-788X
2383-1685
15
1
2009
06
22
NEW SPECIES AND NEW PLANT RECORDS OF LAMIACEAE FROM IRAN
51
56
EN
Ziba
Jamzad
Research Institute of Forests & Rangelands.
<em>Thymus marandensis</em> Jamzad is described as a species new to science. It is closely related to <em>Th. persicus</em> (Ronniger ex Rech. f. ) Jalas,<em>Th. brachychilus</em> Jalas and <em>Th. leucotrichus</em> Hal. <em>Thymus linearis</em> Benth. subsp. <em>linearis</em>, <em>Satureja macrosiphonia</em> Bornm. and <em>Micromeria cristata</em> (Hampe) Griseb. subsp. <em>cristata</em> are reported for the first time from Iran. Their distribution patterns are discussed.
Lamiaceae,Thymus,Satureja,Micromeria,new species,new records,Iran
https://ijb.areeo.ac.ir/article_102724.html
https://ijb.areeo.ac.ir/article_102724_113642ba08d2273423a249a3d5ee70df.pdf
Research Institute of Forests and Rangelands
The Iranian Journal of Botany
1029-788X
2383-1685
15
1
2009
06
22
A NEW RECORD AND A NEW SPECIES OF THE GENUS POA L. (POACEAE) FROM IRAN
57
60
EN
Mahsa
Kavousi
Department of Biology, Islamic Azad University, Campus and Research Branch, Tehran, Iran.
Mostafa
Assadi
Research Institute of Forests and Rangelands.
masadi@rifr-ac.ir
Taher
Nejadsattari
Department of Biology, Islamic Azad University, Campus and Research Branch, Tehran, Iran.
<em>Poa damavandica</em> Assadi & Kavousi is described as a new species from Iran based on a specimen has been collected from high altitudes of Damavand Mountain. It belongs to the section <em>Stenopoa</em> subsection <em>Stenopoa</em>. The new species is compared with <em>Poa araratica, P. sterilis, P. stepposa and</em><em> P. rangkulensis</em> as close affinities. Moreover, <em>Poa glauca</em> is reported as a new record from Iran collected from Mazandaran province.
Poa,Poaceae,new species,Iran
https://ijb.areeo.ac.ir/article_102725.html
https://ijb.areeo.ac.ir/article_102725_b77233b975e5e26803f80d670fc13821.pdf
Research Institute of Forests and Rangelands
The Iranian Journal of Botany
1029-788X
2383-1685
15
1
2009
06
22
TWO NEW RECORDS AND A NEW COMBINATION OF THE GENUS AELUROPUS TRIN (POACEAE) FOR THE FLORA OF IRAN
61
62
EN
Mansooreh
Khodashenas
Organization of Agricultural and Natural Resources Centre, Kerman, Iran.
<em>Aeluropus lagopoides</em> var. <em>mesoptamica</em> is reported from Iran as a new record. This variety is characterized by the following characters: very short and stiff vegetative leaves, short stem and small capitate inflorescence. <em>A. pungens</em> var <em>hirtulus</em> is also reported as another new record for Iran. <em>A. lagopoides</em> var. <em>repens</em> is as a new combination based on <em>A. lagopoides</em> subsp. <em>repens</em>. These two taxa differ from the typical species by having hairy leaves.
Aeluropus,Poaceae,new records,new combination,Iran
https://ijb.areeo.ac.ir/article_102726.html
https://ijb.areeo.ac.ir/article_102726_92edeab6258760fe073984faa20de38a.pdf
Research Institute of Forests and Rangelands
The Iranian Journal of Botany
1029-788X
2383-1685
15
1
2009
06
22
FOUR NEW TAXA AND A NEW RECORD OF PAPAVER L. (PAPAVERACEAE) FOR THE FLORA OF IRAN
63
67
EN
Zahra
Tavakkoli
Department of Biology, Tarbiat Moallem University, Tehran, Iran.
zatavakkoli6@gmail.com
In a taxonomic revision of the genus <em>Papaver </em>in Iran for the preparation of the treatment of Flora of Iran, <em>Papaver dubium </em>subsp. <em>erosum </em>var. <em>album</em> is described as a new variety. It differs from the typical variety by having white petals. <em>Papaver dubium </em>subsp. <em>erosum</em> var. <em>maculatum </em>is another new variety that differs from the typical variety by having black spots at the base of petals.<em> P. decaisnei </em>var. <em>setosum</em> and var. <em>patulum </em>from the sect. <em>Papaver </em>are described as new varieties from C, S, and NE of Iran. They are distinguished from the typical variety by having appressed or spreading seta on the pedicels respectively. <em>Papaver dubium </em>subsp. <em>glabrum </em>is recorded here from Iran (Karaj and Khorassan province) for the first time. For these taxa, morphological characters, identification key and distribution maps are also presented.
Iran,new record,New varieties,Papaver,Taxonomy
https://ijb.areeo.ac.ir/article_102727.html
https://ijb.areeo.ac.ir/article_102727_d5ccf3bd481097c96cd2c621c4d26a91.pdf
Research Institute of Forests and Rangelands
The Iranian Journal of Botany
1029-788X
2383-1685
15
1
2009
06
22
AR(CYPERACEAE), NEW GENUS AND SPECIES RECORDS FROM IRAN THRAXON P. BEAUV (GRAMINEAE) AND CAREX CARYOPHYLLEA
68
71
EN
Bhnam
Hamzeh'ee
Research Institute of Forests and Rangelands.
bhamzeh@yahoo.com
Alireza
Naqinezhad
Department of Biology, Faculty of Science, University of Mazandaran, Babolsar, Mazandaran.
<em>Arthraxon hispidus</em> (Thunb.) Makino var. <em>hispidus</em> (<em>Gramineae</em>) and <em>Carex caryophyllea </em>Latourr. (<em>Cyperaceae</em>) are recorded for the first time as new genus and species for the flora of Iran. These taxa have been collected from Kiashahr (Gilan prov.) and Arasbaran Protected Area (Azerbaijan prov.) in N and NW of Iran. Taxonomical characteristics, localities, habitats, geographical distributions and the closest genus and species are discussed here.
https://ijb.areeo.ac.ir/article_102738.html
https://ijb.areeo.ac.ir/article_102738_becef0e9400e9e023db30281d389a67c.pdf
Research Institute of Forests and Rangelands
The Iranian Journal of Botany
1029-788X
2383-1685
15
1
2009
06
22
PYRUS PASHIA (ROSACEAE), A NEW RECORD FOR THE FLORA OF IRAN
72
75
EN
Asghar
Zamani
Central herbarium of Tehran University, School of Biology, University College of Science, University of Tehran.
asgharzam@khayam.ut.ac.ir
Farideh
Attar
Central herbarium of Tehran University, School of Biology, University College of Science, University of Tehran.
fattar@khayam.ut.ac.ir
Mohammad Reza
Joharchi
Ferdowsi University of Mashhad Herbarium (FUMH), Mashhad, Iran
joharchi@um.ac.ir
<em>Pyrus pashia</em> is reported as a new record from NE Iran. This species is distinguished from its closest relative, <em>P. boissieriana</em>, by having more or less ovate, acuminate leaves and fruit size. An illustration of this species in fruit and flower and a geographical distribution map and also photographs of the two species are given.
Pyrus,Rosaceae,new record,Iran
https://ijb.areeo.ac.ir/article_102741.html
https://ijb.areeo.ac.ir/article_102741_6330a0af062079a08008cc107d94ab74.pdf
Research Institute of Forests and Rangelands
The Iranian Journal of Botany
1029-788X
2383-1685
15
1
2009
06
22
TWO NEW RECORDS OF LOTUS (FABACEAE) AND PHALARIS (POACEAE) FROM IRAN
76
79
EN
Hosein
Maroofi
Research Center of Agriculture and Natural Resources of Kurdistan province.
hosein_maroofi@yahoo.com
Among the materials collected from Kurdistan and West Azerbaijan, two new records to Iran namely <em>Lotus aegaeus</em> and <em>Phalaris canariensis</em> were determined. The species are reported and illustrated in this paper. Descriptions of the species are presented.
Lotus aegaeus,Fabaceae,Phalaris canariensis,Poaceae,new records,Kurdestan,Iran
https://ijb.areeo.ac.ir/article_102745.html
https://ijb.areeo.ac.ir/article_102745_180b411828fc5a2f133c2caa4271b68a.pdf
Research Institute of Forests and Rangelands
The Iranian Journal of Botany
1029-788X
2383-1685
15
1
2009
06
22
A NEW RECORD FOR THE FLORA OF IRAN: MARRUBIUM DUABENSE MURATA (LAMIACEAE)
80
81
EN
Farrokh
Ghahremaninejad
0000-0001-5860-9976
Department of Biology, Faculty of Science, Tarbiat Moallem University, 49 Dr. Mofatteh Avenue, 15614 Tehran, Iran.
ghahremaninejad@gmail.com
Atefeh
Ezazi
Department of Biology, Faculty of Science, Tarbiat Moallem University, 49 Dr. Mofatteh Avenue, 15614 Tehran, Iran.
<em>Marrubium duabense</em> Murata is recorded here for the first time from Iran. This species was known exclusively from Afghanistan; its recording from Iran shows that its distribution is extended to the west to Iran as well. With this record the number of <em>Marrubium</em> species in Iran counts to 11.
Iran,new record,Marrubium duabense
https://ijb.areeo.ac.ir/article_102749.html
https://ijb.areeo.ac.ir/article_102749_b25e9a85ecd9db31c2516dcc3ea704ad.pdf
Research Institute of Forests and Rangelands
The Iranian Journal of Botany
1029-788X
2383-1685
15
1
2009
06
22
TRIGONELLA CAERULEA (FABACEAE), AN AROMATIC PLANT FROM ARDABIL PROVINCE, IRAN
82
84
EN
Mikaeil
Badrzadeh
Department of Natural Resources, College of Agriculture, University of Mohaghegh Ardabili, Ardabil, Iran.
m.badrzadeh@gmail.com
Leili
Ghafarzaseh-namazi
Department of Agronomy, College of Agriculture, the University of Mohaghegh Ardabili, Ardabil, Iran.
<em>Trigonella caerulea</em> (L.) Ser is an annual plant growing as a wild species of <em>Fabaceae</em> family in Iran. This wild fenugreek is described and illustrated for the first time from Ardabil province. In addition, the chromosome numbers of <em>Trigonella caerulea</em> was counted for the first time in Iran. The result showed that the plant is a diploid with 2n = 16.
Trigonella caerulea,Fabaceae,new record,chromosome number,Ardabil,Iran
https://ijb.areeo.ac.ir/article_102753.html
https://ijb.areeo.ac.ir/article_102753_e7cc1f4615c1539e22b191c51a084471.pdf
Research Institute of Forests and Rangelands
The Iranian Journal of Botany
1029-788X
2383-1685
15
1
2009
06
22
BIOSYSTEMATIC STUDY OF ONOBRCHIS VICIIFOLIA SCOP. AND ONOBRCHIS ALTISSIMA GROSSH. (FABACEAE) IN IRAN
85
95
EN
Masoud
Ranjbar
Department of Biology, Herbarium division, Bu-Ali Sina University, P.O.Box 65175/4161, Hamedan, Iran.
ranjbar@ basu.ac.ir
Roya
Karamian
Department of Biology, Herbarium division, Bu-Ali Sina University, P.O.Box 65175/4161, Hamedan, Iran
Anahita
Hadadi
Department of Biology, Herbarium division, Bu-Ali Sina University, P.O.Box 65175/4161, Hamedan, Iran.
Morphologic characters, meiotic chromosome number and behavior were analyzed in different wild populations of <em>Onobrchis</em> <em>viciifolia</em> Scop. and <em>Onobrychis altissima</em> Grossh.belonging to <em>O</em>. sect. <em>Onobrychis</em> in Iran. All Iranian populations of <em>O. viciifolia</em> and<em> O. altissima</em> studied here were tetraploid and showed 2<em>n</em> = 4<em>x</em> = 28 chromosome number. The basic chromosome numbers (<em>x</em> = 7) is consistent with the proposed base number of <em>x</em> = 7 for <em>O</em>. <em>viciifolia </em>and <em>O. altissima</em>. For <em>O. altissima</em>, two ploidy levels (2<em>n</em> = 2<em>x</em> = 14 and 2<em>n</em> = 4<em>x</em> = 28) have been reported from Caucasus. Results support the hypothesis that <em>Onobrchis viciifolia</em> and <em>O. altissima</em> are related as a progenitor-derivative pair of species. The fact that no diploid populations of <em>O. altissima</em> were found in Iran, would suggest a relatively recent origin of the species, most probably in Caucasus. Almost these taxa displayed regular bivalent pairing and chromosome segregation at meiosis. Meiotic abnormalities observed included varied degree of sticky chromosomes with laggards and bridges in anaphase to telophase, asynchronous nuclei in metaphase II and desynapsis in metaphase I. Also <em>O.</em> <em>viciifolia</em> Scop. is considered as a new record from this section for Iran. These taxa of the genus are cultivated as fodder in Iran.
chromosome number,Fabaceae,meiotic behavior,Onobrychis sect. Onobrychis
https://ijb.areeo.ac.ir/article_102755.html
https://ijb.areeo.ac.ir/article_102755_4a137b2148163629b61babb318b5d5f4.pdf
Research Institute of Forests and Rangelands
The Iranian Journal of Botany
1029-788X
2383-1685
15
1
2009
06
22
ANATOMICAL STUDIES ON FOURTEEN SPECIES OF THE GENUS COTONEASTER L. (ROSACEAE) IN IRAN
96
104
EN
Nematollah
Raei Niaki
Central herbarium of University of Tehran, School of Biology, University College of Science, University of Tehran, P. O. Box 14155- 6455, Tehran, Iran.
Farideh
Attar
Central herbarium of University of Tehran, School of Biology, University College of Science, University of Tehran, P. O. Box 14155- 6455, Tehran, Iran.
fattar@khayam.ut.ac.ir
Hosein
Maroofi
Research Center of Agricultural and Natural Resources of Kurdistan, P. O. Box 66169- 36311- 714, Kurdestan, Iran.
hosein_maroofi@yahoo.com
In this study, anatomical features of blade and midrib region in 14 species of genus <em>Cotoneaster</em> belonging to two sections <em>Cotoneaster </em>and <em>Chaenopetalum</em> are presented. These species include <em>Cotoneaster nummularioides</em>, <em>C. kotschyi, C. turcomanicus, C. multiflorus, C. nummularius, C. hissaricus, C. luristanicus, C. suavis, C. morulus, C. ovatus, C. discolor, C. assadii, C. melanocarpus </em>and <em>C. persicus</em>. Among 30 studied characters, five characters (quantitative features such as presence of fiber, presence of cuticle and external xylem) are constant in different species. Twenty four characters 24 are variable in several species and several individuals of the same species. Only one character (i.e. number of palisade parenchyma layer) was variable and useful in separation of species.
Cotoneaster,Rosaceae,Anatomy,Taxonomy,Iran
https://ijb.areeo.ac.ir/article_102758.html
https://ijb.areeo.ac.ir/article_102758_72d68cf7e25241ba7a6e1f5d0968f2cb.pdf
Research Institute of Forests and Rangelands
The Iranian Journal of Botany
1029-788X
2383-1685
15
1
2009
06
22
MICROMORPHOLOGICAL AND ANATOMICAL STUDIES OF CERTAIN SPECIES OF VERBASCUM (SCROPHULARIACEAE) IN WEST AZERBAIJAN, IRAN
105
113
EN
Somayyeh
Kheiri
Department of Biology, Faculty of Science, Urmia University, P.O. Box: 57135-165, Urmia, Iran.
somayyeh.kh@gmail.com
Masoud
Khayami
Department of Biology, Faculty of Science, Urmia University, P.O. Box: 57135-165, Urmia, Iran.
Ahmad
Mahmoodzadeh
Department of Biology, Faculty of Science, Urmia University, P.O. Box: 57135-165, Urmia, Iran.
The high morphological diversity among the species of the genus <em>Verbascum</em> causes problems in the delimitation of the species. Therefore, there is a difficulty in offering a natural and well organized classification for the species of the genus. This research was based on the leaf, seed coat and fruit anatomy, and seed micromorphology of the species growing in Iran. The shape and density rate of epicarpic idioblasts can be a determining character for <em>Verbascum</em> species.Two types of mesophyll were found within the species: Dorsiventral and Isobilateral. Also two types of thickness ratio of endocarp to pericarp (2/3, 1/2) were distinguished. However, a slight difference has been observed in the shape of epidermal cells of seeds and the presence of vesicle on the lateral walls of the epidermal cells of seeds.
Verbascum,Iran,seed,micromorphology,leaf,Fruit,Anatomy,Scrophulariaceae
https://ijb.areeo.ac.ir/article_102759.html
https://ijb.areeo.ac.ir/article_102759_421f6afbdf1ae71d399f3567fc8f3332.pdf
Research Institute of Forests and Rangelands
The Iranian Journal of Botany
1029-788X
2383-1685
15
1
2009
06
22
STUDIES OF POLLEN GRAINS IN THE SECT. STENOCEPHALAE (COUSINIA CASS.-ASTERACEAE) IN IRAN
114
128
EN
Aghdas
Saber
Central Herbarium of Tehran Uuniversity, School of Biology, College of Sciences, University of Tehran.
Farideh
Attar
Central Herbarium of Tehran Uuniversity, School of Biology, College of Sciences, University of Tehran.
fattar@khayam.ut.ac.ir
Seyyedeh Bahereh
javadi
Department of Botany, Iranian Research Institute of Plant Protection, P.O. Box 1454, Tehran 19395, Iran.
sb_djavadi@hotmail.com
Pollen morphology of 25 species of section <em>Stenocephalae</em> Bunge from genus <em>Cousinia</em> Cass. is described and compared using Light and Scanning Electron Microscope. The pollen grains are prolate, isopolar , tricolporate and exine is densely or loosely verrucate. Based on morphological characters, species of this section can be divided to two groups: 1. species with narrow cylindrical heads and leathery leaves, 2. species with ovate heads and herbaceous leaves. The pollen grains of the first group with more than 20 verrucae and the second group with 3-30 verrucae in each 25 µm² of pollen surface are differentiated from each other. Indeed, these two detached grouping for species of sect. <em>Stenocephalae</em> are somewhat supporting each other.
Cousinia,sect. Stenocephalae,Pollen morphology,Iran
https://ijb.areeo.ac.ir/article_102762.html
https://ijb.areeo.ac.ir/article_102762_77d92786fee5b6803b466806ac9888dc.pdf
Research Institute of Forests and Rangelands
The Iranian Journal of Botany
1029-788X
2383-1685
15
1
2009
06
22
MORPHOLOGICAL AND MICROMORPHOLOGICAL STUDIES OF NONEA (BORAGINACEAE: TRIBE BORAGINEAE) IN IRAN
129
139
EN
Manizheh
Pakravan
Department of Biology, Faculty of Science, Alzahra University, Tehran, Iran.
mpf176@yahoo.com
Atiye
Nejhad Falatoury
Department of Biology, Faculty of Science, Alzahra University, Tehran, Iran.
anfmk@yahoo.com
Ahktar
Tavassoli
Department of Biology, Faculty of Science, Alzahra University, Tehran, Iran.
tavastars@yahoo.com
<em>Nonea</em> Medik. is one of the major genera of <em>Boraginaceae</em> tribe <em>Boragineae</em>, with approximately 35 species in the world. In this study we focused on reproductive characters, which are not generally affected by environment. Our selected reproductive characters have been the stigma form, position of stigma towards the anthers, form of faucal scales, color, form and symmetry of flowers and characters of mericarps. An identification key using morphological and micromorphological characters has also been proposed. In the present paper above mentioned characters have been described for the first time in <em>N. turcomanica </em>M. Pop<em>, N. lutea</em> (Desr.) DC. var<em>. flavescens</em> Lipsky<em>, N. persica </em>Boiss. var.<em> suchtelenioides</em> (Riedl) Falatoury & Pakravan, <em>N. diffusa </em>Boiss. et Buhse, <em>N. rosea </em>(Bieb.) Link and<em> N. melanocarpa </em>Boiss.<em> Nonea suchtelenioides</em> H. Riedl has been decreased to the variety level of <em>Nonea persica </em>as<em> N. persica </em>Boiss. var.<em> suchtelenioides</em> (Riedl) Falatoury & Pakravan.
Nonea,Boraginaceae,Taxonomy,micromorphology,Stigma,faucal scale,Iran
https://ijb.areeo.ac.ir/article_102763.html
https://ijb.areeo.ac.ir/article_102763_77f8f2d141a82af7cde9c0d02c439e9d.pdf